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solute concentration of urine is equal to the solute
concentration of the body fluids.
Water conservation
by mammalian
kidneys
Related topics Solar radiation and climate(C1) Plants and water (D2)
Microclimate (C2) Ecosystem patters (S1)
The properties of water (D1)
E1 温度与代谢
要 点
恒温动物和变
温动物
按照有机体的体温调节过程有可能对有机体进行分类。
一种可能的划分是恒温动物和变温动物。当环境温度
升高时,恒温动物保持着大致恒定的体温,而变温动
物的体温随环境温度而改变。这种分类的一个问题是,
即使是典型的恒温动物也经历了降低体温的时期。在
有机体之间,可供选择的另一种划分描述如下。
E1 TEMPERATURE AND
METABOLISM
It is possible to categorize organisms according to their
temperature regulation processes,One possible division
is between homeotherms and poikilotherms,As
environmental temperature rises,homeotherms maintain
an approximately constant body temperature,while the
body temperature of poikilotherms varies with
environmental temperature,One problem with this
classification is that even classic homeotherms
experience periods of reduced temperature.,an
alternative distinction between organisms is described
below.
Key Notes
Homeotherms and
poikilotherms
外温动物和内
温动物
像爬行动物等有机体属外温动物,它们很大程度地依
赖于外来的热来提高自己的体温。内温动物是能够在
体内产热以升高体温的有机体。鸟和哺乳动物属于这
一类。 热中性区 ( thermoneutral zone)是环境的温度范
围,在这个温度范围内,内温动物仅使用一个最小的
代谢率以维持恒定的体温。环境温度离热中性区越远,
内温动物维持恒定体温消耗的能量越多。
热 交 换 所有的有机体从它们的环境中得到热,也将产生的热
散失到它们的环境中。种种生理学和行为学的机理被
用于调节体热。尽管有这些机理,外温性动物的体温
还是明显地随环境条件而改变。
Ectotherms are organisms such as plants,reptiles and
protista which are largely reliant on external sources of
heat to raise their body temperature.endotherms are
organisms capable of generating heat internally in order to
raise their body temperature,Birds and mammals make up
this group,The thermoneutral zone is the range of
enmironmental temperatures in which an endotherm has
only to exert a minimum metabolic effort in order to
maintain a constant body temperature,The further away
from the thermoneutral zone that the environmental
temperature moves,the more energy the endotherm has to
expend to maintain body temperature,
Ectotherms and
endotherms
All organisms gain heat from and lose heat to their
environment as well as producing heat,A variety of
physiological and behavioral mechanisms are used to
regulate heat,Despite these mechanisms,the body
temperature of an ectotherm varies significantly with
environmental conditions.
Heat exchange
温 度 阈 高温可能导致酶失活或代谢组分不平衡,例如植物的
呼吸作用快于光合作用而导致死亡。然而高温对外温
动物最普遍的影响是引起脱水。所有陆生外温动物必
须保持水,但在高温下失水率能够成为致死因子。不
同物种对低温的耐受性有很大的差异,这与结冰、寒
冷和坚硬的过程有关。温度低于 -1℃ 时很多物种被冻死,
这是由于细胞内冰晶形成的损伤效应;那些生活在整
个冰冻冬天的种,经常是处于一种有抵抗力的休眠时
期,这是它们生命周期中的一个阶段。
相关题目 太阳辐射与气候( C1) 对温度的响应( E2)
植物与水( D2) 温度与物种分布( E3)
动物与水( D3) 生态系统格局( S1)
High temperatures may lead to enzyme inactivation or the
unbalancing of components of metabolism; for example,in
plants,respiration may proceed faster than photosynthesis,
leading to death,however,the most frequent effect of high
temperature on ectotherms is dehydration,All terrestrial
ectotherms must conserve water but at high temperatures
rates of water loss can be lethal,There are large differences
between the low temperature tolerances of differing species,
associated with the processes of freezing,chilling and
hardening,Many are killed by temperatures below –1℃
due to the damaging effects of ice-crystal formation within
cells; those that live through freezing winters often do so at
a resistant,dormant stage of their life cycle.
Temperature
thresholds
Related topics Solar radiation and climate(C1) Responses to temperature
(E2)
Plants and water (D2) Temperature and species
Animals and water (D3) distribution (E3)
Ecosystem patterns (S1)
E2 对温度的响应
要 点
温度和酶反应
速度
酶催化反应的速度随温度而增加。在外温动物中这意
味着在较高的温度中代谢活力更快。温度系数( Q 10)
是温度升高 10℃ 对代谢速度影响的指数,经常大约为
2.0。
发育和生长的
速度
在非致死温度范围内,温度对外温动物最大的影响很
可能是影响了发育和生长的速度。当用发育速度对体
温做图时,存在着一个直线相关的温度范围。“生理
时间”是将温度和时间相结合的一个测量,应用于外
温动物,它反应了这些有机体的生长和发育依赖于温
度及时间。
The rate of an enzyme catalyzed reaction increases with temperature,
In ectotherms this means that metabolic activity will be faster at
higher environmental temperatures,The temperature coefficient (Q10)
is an indexof the effect of a 10℃ temperature rise on metabolic rate,
and is often near 2.0,
Key Notes
Within the nonlethal temperature range the most important effect
on ectotherms of temperature is likely to be its effect on rate of
development and growth,When rate of development is plotted
against body temperature there exists an extended range of
temperatures over which the relationship is linear,?Physiological
time‘ is a measurement combining temperature and time and
applied to ectothermic organisms,which reflects that growth and
development of these organisms is dependent on environmental
temperature as well as time.
Temperature and
rates of enzyme
reaction
Rates of development
and growth
E2 RESPONSES TO TEMPERATURE
驯化和气候
驯化
温度能够作为一种刺激物起作用,决定有机体是否将
开始发育。春化法是通过低温诱导开花。有机体在实
验室里暴露到较高(或较低)的温度能够改变有机体
的温度反应。有机体对实验环境条件变化产生的适应
性反应称为 驯化 ( acclimation)。有机体对自然环境条
件变化产生的生理适应性反应称为 气候驯化
( acclimatization)。
相关主题 太阳辐射与气候( C1) 温度与物种分布( E3)
温度与代谢( E1) 生态系统格局( S1)
Temperature may also act as a stimulus,determining whether the
organisms will begin development,Vernalization is the induction
of flowering by low temperatures,Exposure of an organism to
higher (or lower) temperatuers in the laboratory can alter the
organisms temperature response,The habituation of an organism‘s
response to changes in laboratory environmental conditions is
termed acclimation,Acclimatization is the habituation of an
organism‘s physiological response to changes in natural
environmental conditions.
Acclimation and
acclimatization
Related topics Solar radiation and climate(C1) Responses to temperature (E2)
Plants and water (D2) Temperature and species
Temperature and metabolism (E1) distribution (E3)
Ecosystem patterns (S1)
The rate of photosynthesis is a gross measurement of the rate at
which a plant captures radiant energy and fixes it into carbon
compounds,Net assimilation is the difference between
photosynthetic assimilation and losses due to respiration,
Therefore,net assimilation will be negative in the dark and will
increase with increasing PAR,The intensity of PAR at which the
gain in photosynthesis equals the losses is known as the
compensation point.
Measurement of
photosynthesis
E3 温度与物种分布
要 点
物种分布与
温度
地球上主要生物群系的分布反应了主要的温度带。然
而当考虑物种分布的时候,很难归结于温度的作用。
在某些情况下,一个物种的分布界限能够归结于阻止
物种生存的致死温度。然而,种的分布限与等温线间
相一致,是一种更广泛分布的相关类型。等温线是在
地图上把具有相同平均温度的地方连结起来的线。总
的说来,一个有机体的分布界限不是由致死温度所决
定,而是由造成竞争强弱条件所决定。
The distribution of the major biomes over the Earth reflects the major
temperature zones,However,it is more difficult to attribute a role to
temperature when considering species distribution,In certain cases
the distribution limits of a species can be attributed to a lethal
temperature which precludes the species‘ existence,However,a more
widespread type of relationship is one which has a correspondence
between the distribution limits of a species and an isotherm,An
isotherm is a line on a map that joins locations having the same mean
temperatures,Overall,an organism‘s limit to distribution is
determined not by lethal temperatures,but by conditions that make it
a poor competitor,
Key Notes
Species distribution
and temperature
E3 TEMPERATURE AND SPECIES
DISTERIBUTION
对温度的进化
反应
温度对个体的影响被进化的变异减弱。阿伦法则
( Allen‘s rule)陈述了来自冷气候中的内温动物与来自
温暖气候的内温动物相比,趋向于具有更短的末端
(耳朵和四肢);因此降低了它们的表面积对体积的
比率。这个规律有很广的适用性。贝格曼规律
( Bergmann‘s rule)讲述了寒冷地区的哺乳动物比温暖
地区的哺乳动物个体趋向于更大,也减低了它们的表
面积与体积的比率。这个规律没有广泛的种间的适用
性,是由于在测定体重中牵涉到其它的重要因子,但
是在种内经常是真实的。
相关主题 温度与代谢( E1) 竞争的性质( I1)
对温度的响应( E2) 生态系统格局( S1)
The effects of temperature on individuals may be moderated by
evolved differences,Allen‘s rule states that endothermic animals
from cold climates tend to have shorter extremities (ears and legs)
compared with animals from warmer climates,thus reducing their
surface area,volume ratio,This rule has widespread applicability,
Bergmann‘s rule states that mammals tend to be larger in colder
areas than warm climates,again to reduce their surface area,
volume ratio,This rule does not have widespread interspecific
applicability due to other important factors involved in
determining body weight,but is often true intraspecifically.
Evolved response to
temperature
Related topics Temperature and metabolism (E1) The nature of competition (I1)Responses to temperature (E2) Ecosystem patterns (S1)
F1 太阳辐射与植物
要 点
辐射能和光合
作用
绿色植物能够利用的惟一能源是辐射能。当叶子截获
辐射能时,它能被吸收、反射或者透射。吸收的部分
能量到达叶绿体,引发了光合作用,在这个过程中,
辐射能被用于转化水和二氧化碳成为糖。太阳辐射包
含了不同波长的光谱。然而,仅有一个有限的光谱带
对光合作用是有效的。这就是光合活性辐射( PAR)带,
对绿化植物是位于 380nm到 700nm之间。
F1 SOLAR RADIATION AND
PLANTS
Radiant energy is the sole energy source that can be
used by green plants,When a leaf intercepts radiant
energy it may be absorbed,reflected or transmitted,
Part of the fraction absorbed reaches the chloroplast,
fuelling photosynthesis,the process where radiant
energy is used to convert water and CO2 into sugars,
Solar radiation contains a spectrum of different
wavelengths,However,only a restricted band of this
spectrum is effective for photosynthesis,This is the
band of photosynthetically active radiation (PAR)
and for green plants lies between 380 and 710 nm..
Key Notes
Radiant energy and
photosynthesis
辐射能转换
效率
计算光合作用效率是有可能的。作为生物化学过程,
光合作用是高效率的;进入反应部位的可用辐射能的
35%转化为潜在能量。在植物水平上的真实效率在
0.5%和 3%之间变化,取决于植物和环境。
C3和 C4植物 植物光合能力中的主要差别是在 C3和 C4植物之间。 C4
植物能捕获 CO2,伴随着水的利用效率比 C3植物更大,
而这优点需要消耗能量。在 C4植物中,光合作用率随
光强度而增加,而 C3植物随光强度增加光合作用渐渐
减小。
It is possible to calculate the efficiency of
photosynthesis,As a biochemical process
photosynthesis is efficient; 35% of usable4 radiant
energy entering a reaction site is converted to
potential energy,The actual efficiency at the plant
level varies between 0.5% and 3.0%,depending on
the plant and the environment.
Efficiency of radiant
energy conversion
A major difference in the photosynthetic capacity of
plants is that between C3and C4 plants,C4 plants are
able to capture CO2 with greater water use efficiency
than C3 plants,but this advantage comes at an energy
cost,In C4 plants the rate of photosynthesis increases
with light intensity,whilst photosynthesis tails off
with increasing light intensity in C3 plants,
C3 and C4 plants
辐射强度的
变化
植物很难获得它们完全的光合作用潜能,是由于水短
缺和辐射强度的改变。光强度的系统变化是太阳辐射
的日节律和年节律。光强度中极少部分的系统变化是
因叶子彼此的相对位置引起。
植物对辐射的
战略和战术
响应
植物种间对辐射强度反应的主要战略差异显示为“阳
地种”和“阴地种”,它们分别具有适应于高的和低
的光辐射范围。同样,植物能够在不同光条件下生长
不同的叶子,作为对光环境的部分战术反应。这一点
最清楚地在单株植物叶冠内的阳叶和阴叶的结构上看
到。
Plants rarely achieve their full photosynthetic potential,
due to water shortage and to variation in the intensity
of radiation,The systematic variations in light
intensity are the diurnal and annual rhythms of solar
radiation,Less systematic variations in light intensity
are caused by the posit