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Rule 2 Ecology is only understandable in the light of evolution.
The huge diversity of organisms,and the wealth of variety in
their morphologies,physiologies and behavior are all the
result of many millions of years of evolution,Thos
evolutionary history has left an indelible impression on each
and every individual,It is only possible to make sense of the
patterns we find today in the light of this evolutionary legacy.
Rule 3 Nothing happens¡¯ for the good of the species¡¯.
A very common misconception is the idea that patterns of
behavior in organisms which appear to be costly to an
individual occur¡® for the good of the species¡®,This is
absolutely and completely wrong,Natural selection will
favor those genes which are passed on to the most offspring,
even if these genes may cause a reduction in the
species¡®population size.
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Rule 4 Genes and environment are both important,
The environment an organism finds itself in plays an
important role in determining the options open to that
individual,The genes which define an organism¡®s makeup
are also of fundamental importance,To understand ecology
it is important to appreciate the fundamental nature of both
of these factors and the fact that they interact.
Rule 5 Understanding complexity requires models.
Ecology is a complex subject,with huge variation at
almost every scale-millions of species,each with
considerable genetic variation,varying numbers and
ever-changing behaviors in a complex and dynamic
environment,To understand it,it is necessary to clearly
identify specific questions and then formulate hypotheses
which can be tested,It is often very useful to frame the
hypothesis in mathematical terms to avoid ambiguity and
confusion which are often inevitable in a verbal model,
Mathematical models are widely used in ecology.
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Rule 6 ¡®Story-telling¡¯ is dangerous.
In attempting to explain ecological patterns or
relationships,it is easy to slip into a make-believe
world where every observation is readily explained by
some ad hoc assertion ¨C?story-telling¡®,The temptation
to advance hypotheses as facts should be avoided at all
costs.
Rule 7 There are hierarchies of explanations.
For any observation there is often an immediate cause
that can be diagnosed,Often this causal explanation is
insufficiently informative and we need to probe deeper
to reach a fuller grasp of the situation,Even if a
phenomenon is ?explained¡® there may well be further
and deeper explanations which allow us to see the
fuller picture.
¹æÂÉ 8£ºÓлúÌå
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Rule 8 There are multiple constraints on organisms.
Whilst the total diversity of form,function and
environmental resilience exhibited by organisms is awe-
inspiring,each individual (and,to a slightly lesser
extent,each species) operates within a relatively narrow
range of constraints,Constraints fundamentally take
two forms,(i) physical and (ii) evolutionary,Evolution
cab never reach ?perfection¡® because of these
constraints and organisms are essentially hotchpotches
of numerous compromises,
¹æÂÉ 9£º»ú»áÊÇ
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Rule 9 Chance is important.
Chance events play a critical role in ecology,The
opening of a gap in a forest canopy or the breaching of
a sand dunce after a storm will have a major impact on
the ecology of the local fauna and flora,but both are
unpredictable in either time or location,The
importance of chance events in ecology does not mean
ecological patterns are wholly unpredictable,but it
necessarily places boundaries on the potential level of
predictive detail.
Rule 10 There boundaries of ecology are in the mind of the ecologist.
Ecology is a broad science,covering both organisms
and physical environments and hence excludes little as
potentially relevant,Mathematics,chemistry and
physics are tools essential to the understanding of
ecology.
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B1 ADAPTATION
Key Notes
Fitness is measure of the ability of an individual to produce
viable offspring and contribute to future generations,
Individuals vary in their relative fitness,and this variation is
due partly to genetic differences among individuals and
partly to environmental influences.
The individuals in a species which have the highest fitness will
contribute disproportionately to the subsequent generations,If
fitness differences have a genetic component,then the genetic
make-up of the subsequent generations will be altered,This
process is known as natural selection or ?survival of the fittest¡®.
Fitness
Natural selection
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Any heritable trait possessed by an organism which aids
survival or reproduction is an adaptation,Such traits may
be physiological,morphological or behavioral,Adaptation
is the result of natural selection.
The genotype is the genetic composition of an individual,
The phenotype is the individual organism,a product of the
interaction between its genotype environmental influences
on its genotype is known as phenotypic plasticity (e.g,
human suntan,wind-shaped plants locust morph (solitary
or migratory)).
Related topics Ten rules in ecology (A2) Speciation (O2)
Genetic Variation (O1)
Adaptation
Genotype and
phenotype
B2 Ó¦¸¶»·¾³±äÒì
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Ìõ ¼þ ÒýÆðÓлúÌå¶ÔÆä·´Ó¦µÄ, ¿É±äµÄ»·¾³Òò×ÓÊÇÌõ¼þ
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Key Notes
Variable environmental factors which organisms respond to
are conditions,Examples include temperature,acidity and
salinity,Conditions are not depletable ¨C they are not used up
or consumed by an organism.
Anything which the organism uses up or depletes is a
resource for that organism,Thus,nectar is a resource for
honey bees,and light is a resource for all green plants.
B2 COPING WITH ENVIRONMENTAL
VARIATION
Conditions
Resources
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Most organisms have to cope with a continually changing
external environment over a range of timescales,Some
environmental factors may change over seconds or
minutes (e.g,sunlight intensity when,there is patchy cloud)
whilst others may change daily or seasonally or over a
much change daily or seasonally or over a much longer
period (e.g,glaciation cycles).
Environmental
variation
Biological cells cannot function with a wildly
fluctuating environment and organisms therefore take
steps to limit their internal variation.
Internal
regulation
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The maintenance of a relatively constant internal
environment by an organism in a variable external
environment is called homeostasis,All organisms adopt
a degree of homeostatic control,but,as t rule,large
organisms are more decoupled from their external
environment than small ones.
Most biological homeostatic mechanisms act in a
broadly similar way,if the current internal level of a
factor(e.g.temperature or osmolarity) is too high,the
mechanism will reduce it; if the level is too low the
mechanism will increase it,This process,by which the
response is opposite to the signal,is known as negative
feedback.
Homeostasis
Negative feedback
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Organisms can cope with variation in their external
environment (though different species may differ markedly),
The upper and lower extremes of species limits of tolerance,
Usually,growth will not occur at these extremes but in a
narrower range of conditions,and fitness will be greatest
only for a yet narrower optimal range.
Plants and water (D2) responses to
temperature (E2)
Animals and water (D3) solar radiation and
plants (F1)
Related topics
tolerance
B3 Éú ̬ λ
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The ecological niche of an organism is the position it
fills in its environment,comprising the conditions
under which it is found,the resources it utilizes and
the time it occurs there.
Key Notes
The habitat of an organism is the physical
environment it is found in,for example,a temperate
broad-leaved woodland,Each habitat provides.
Each condition or resource which defines the niche
of an organism contributes one dimension to the
space in which the organism¡®s niche,and is the
multidimensional niche space,or¡®n-dimensional
hypervolume¡®.
B3 THE NICHE
Niche
Habitat
Multidimensional
niche space
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The niche space an organism can fill in the absence of
competition or predation is known as the fundamental
niche.
The niche space occupied by an organism when
competition and predation occur is the realized niche,
which is always a subset of the fundamental niche.
The nature of competition (11) Resource
partitioning(13)
Intraspecific competition (12)
Fundamental niche
Realized niche
Related topics
C1 Ì«Ñô·øÉäÓëÆøºò
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Key Notes
Solar energy drives climatic processes,Energy from the
sun strikes the Earth where high-energy wavelengths of
light are absorbed and re-emitted in the form of radiant
heat,Warmed air rises,a